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Recent Papers

Review Paper: Hoseini, M., Arzani, A. (2023). Epigenetic adaptation to drought and salinity in crop plants. J. Plant Mol. Breed 11 (2): 1-16. 

Mirniyam G., Rahimmalek M., Arzani A.,Yavari P., Sabzalian M.R., Ehtemam M.H., Szumny A. (2023). Phytochemical, morphological, and physiological variation in different ajowan (Trachyspermum ammi L.) populations as affected by salt stress, genotype × year interaction and pollination system. Int. J. Mol. Sci. 24, 10438.

Haghighi, R., B.E. Sayed-Tabatabaei, SAM. Mirmohammady Maibody, A. Arzani, M. Talebi (2023) Expression of flowering repressor gene CsSVP, carbohydrates, and antioxidants affected by plant growth regulators in saffron. J. Plant Growth Regul. 42: 1171–1185,

Review Paper: Soleimani, M., A, Arzani, V. Arzani, T.H. Roberts (2022) Phenolic compounds and antimicrobial properties of mint and thyme (Lamiaceae, tribe Mentheae). J. Herb. Med. 36: 100604, doi: 10.1016/j.hermed.2022.100604.

Ebrahim F, Arzani A, Rahimmalek M, Rezaei S, and Cheghamirza K (2022) Yield related traits and tolerance indices to screen salinity tolerant genotypes in cultivated and wild barley. Gesunde Pflanzen 74: 1099–1108. doi: 10.1007/s10343-022-00692-5

Mahdavi S, Arzani A, Mirmohammady Maibody SAM and Kadivar M (2022) Grain and flour quality of wheat genotypes grown under heat stress. Saudi J. Biol. Sci. 29:103417. doi:10.1016/j.sjbs.2022.103417.

Mirniyam G, Rahimmalek M, Arzani A, Matkowski A, Gharibi S, Szumny A (2022). Changes in essential oil composition, polyphenolic compounds and antioxidant capacity of ajowan (Trachyspermum ammi L.) populations in response to water deficit. Foods 11, 3084.

Review Paper: Brouns F., S. Geisslitz, C. Guzman, T.M. Ikeda, A. Arzani, G. Latella, S. Simsek, M. Colomba, A. Gregorini, V. Zevallos, V. Lullien-Pellerin, D. Jonkers, P.R. Shewry (2022). Do ancient wheats contain less gluten than modern bread wheat, in favour of better health? Nutrition Bulletin 47 (2). doi: 10.1111/nbu.12551.

Gharaghanipor N, Arzani A, Rahimmalek M and Ravash R (2022) Physiological and transcriptome indicators of salt tolerance in wild and cultivated barley. Front. Plant Sci. 13:819282. doi: 10.3389/fpls.2022.819282

Omrani S., A. Arzani, M. Esmaeilzadeh Moghaddam, M. Mahlooji (2022) Genetic analysis of salinity tolerance in wheat (Triticum aestivum L.). PLoS ONE 17(3): e0265520.

Mortazavi S.E. A. Mirlohi, A. Arzani (2022). Overexpression of choline oxidase gene in three filial generations of rice transgenic lines. Iran J. Biotechnol. 20 (2): 89-98. doi: 10.30498/IJB.2022.245350.2850

Chavoushi M., M. Kadivar, A. Arzani, M.R. Sabzalian (2022). Relationships between grain, flour, and dough quality characteristics and solvent retention capacity tests of twelve triticale cultivars and parental species. Food Chem. 371: 131283, doi:10.1016/j.foodchem.2021.131283

Kiani R.,  A. Arzani, S.A.M. Mirmohammady Maibody (2021) Polyphenols, flavonoids and antioxidant activity involved in salt tolerance in wheat, Aegilops cylindrica and their amphidiploids. Front. Plant Sci. 12:646221 | doi: 10.3389/fpls.2021.646221.

Bahrami F., A. Arzani, M. Rahimmalek (2021) Tolerance to high-temperature at reproductive stage: trade-offs between phenology, grain yield and yield-related traits in wild and cultivated barleys. Plant Breed. 140 (5): 812-826, doi: 10.1111/pbr.12953

Kiani R., A. Arzani, S. A. M. Mirmohammady Meibody, M. Rahimmalek, K.Razavi (2021) Morpho-physiological and gene expression responses of wheat by Aegilops cylindrica amphidiploids to salt stress. Plant Cell Tiss. Org. Cult. 144 (3): 619–639.  doi: 10.1007/s11240-020-01983-3.

Mahdavi S., A. Arzani, S.A.M. Mirmohammady Maibody, A.A.Mehrabi  (2021) Photosynthetic and yield performance of wheat (Triticum aestivum L.) under sowing in hot environment. Acta Physiol. Plant. 43: 106. doi:10.1007/s11738-021-03278-2.

Kiani, R., A, Arzani, S.A.M. Mirmohammady Maibody, M. Rahimmalek, T. Ayers (2021) Hybridization of wheat and Aegilops cylindrica: development, karyomorphology, DNA barcoding and salt tolerance of the amphidiploids. J. Plant Biochem. Biotechnol. 30: 943–959; doi: 10.1007/s13562-021-00694-w.

Tohidi B., M. Rahimmalek, A. Arzani (2021) Variation in phytochemical, morphological, and ploidy levels of Iranian Thymus species. Chem. Biodiv. 18: doi:10.1002/cbdv.202000911.

Amiri R., S. Bahraminejad, K. Cheghamirza, A. Arzani (2020) Genetic analysis of iron and zinc concentrations in bread wheat grains. Journal of Cereal Science 95: 103077.

Hodaei M., M. Rahimmalek, A. Arzani (2020) Variation in bioactive compounds, antioxidant and antibacterial activity of Iranian Chrysanthemum morifolium cultivars and determination of major polyphenolic compounds based on HPLC analysis. J Food Sci Technol.

Tohidi B., M. Rahimmalek, A. Arzani and M.R. Sabzalian (2020) Thymol and carvacrol, phenolics and antioxidant accumulation in Thymus species in response to different light spectra. Food Chem. 307: 125521, doi:10.1016/j.foodchem.2019.125521

Tohidi B., M. Rahimmalek, A. Arzani and H. Trindade (2020) Sequencing and variation of terpene synthase gene (TPS2) as the major gene in biosynthesis of thymol in different Thymus species. Phytochemistry 169: 112126.

Ebrahim, F., A. Arzani and M. Rahimmalek (2020) Salinity tolerance of wild barley Hordeum vulgare ssp. spontaneum. Plant Breeding 139: 9:304–316, doi: 10.1111/pbr.12770.

 Ariño-Estrada G., G. Mitchell , P. Saha , A. Arzani , S. Cherry , E. Blumwald , A. Kyme (2019) Imaging salt uptake dynamics in plants using PET. Sci. Rep. 9, Article number: 18626.

Akrami M., A. Arzani (2019) Inheritance of fruit yield and quality in melon (Cucumis melo L.) grown under field salinity stress. Sci Rep. 9.

Hodaei M., M. Rahimmalek and A. Arzani (2019) Genetic diversity of Iranian Chrysanthemum morifolium cultivars using morphological traits and sequence‑related amplified polymorphism (SRAP) markers. Hort. Environ. Biotechnol. 60: 753–765.

Arabbeigi M., A. Arzani, M.M. Majidi (2019) Expression profiles of P5CS and DREB2 genes under salt stress in Aegilops cylindrica. Russ. J. Plant Physiol. 66 (4): 583–590.

Abbasi Z., A. Arzani, M.M. Majidi, J. Amirhooshang, A. Rajabi (2019) Genetic analysis of sugar yield and physiological traits in sugar beet under salinity stress conditions. Euphytica 215: 99. doi: 10.1007/s10681-019-2422-5

Review Paper: Mohammadinejad R., Shavandi A., Raie D.S., Sangeetha J., Soleimani M., Shokrian Hajibehzad S., Thangadurai D., Hospet R., Popoola J.O., Arzani A., Gómez-Lim M.A., Iravani S. and Varma R.S. (2019) Plant molecular farming: Production of metallic nanoparticles and therapeutic proteins using green factories. Green Chem. 21: 1845-1865.

Bahrami F., A. Arzani and M. Rahimmalek (2019) Photosynthetic and yield performance of wild barley (Hordeum vulgare ssp. spontaneum) under terminal heat stress. Photosynthetica 57 (1): 9-17. DOI:  10.32615/ps.2019.009 

Book Chapter: Arzani A. (2019) Emmer (Triticum turgidum  ssp. dicoccum) flour and bread. In: Preedy V.R., R.R. Watson, and V.B. Patel (eds.) Flour and fortification in health and disease prevention. 2nd edition. Elsevier, DOI: 10.1016/B978-0-12-814639-2.00007-1

Book Chapter: Arzani A. (2018) Engineering programmed cell death pathways for enhancing salinity tolerance in crops. In: Kumar V., Wani S.H., Suprasanna P. and Tran LPS. (eds.) Salinity responses and tolerance in plants, Volume 2. Springer. DOI: 10.1007/978-3-319-90318-7_5

Salehi M., A. Arzani, M. Talebi, and A. Rokhzadi (2018) Genetic diversity of wheat wild relatives using ssr markers. Genetika 50: 131-141.

Arabbeigi M., A. Arzani, M.M. Majidi1, B.E. Sayed-Tabatabaei, P. Saha (2018)  Expression pattern of salt tolerance-related genes in Aegilops cylindricaPhysiol. Mol. Biol. Plant. 24: 61-73. DOI

Hodaei M., M. Rahimmalek, A. Arzani and M. Talebi (2018). The effect of water stress on phytochemical accumulation, bioactive compounds and expression of key genes involved in flavonoid biosynthesis in Chrysanthemum morifolium L. Ind. Crops Prod. 120: 295–304.

Akrami, M. and A. Arzani (2018) Physiological alterations due to field salinity stress in melon (Cucumis melo L.) Acta Physiol. Plant. 40: 91 DOI: 10.1007/s11738-018-2657-0

REVIEW PAPER: Arzani A. and M. Ashraf (2017) Cultivated ancient wheats (Triticum spp.): A potential source of health-beneficial food products. Compr. Rev. Food Sci. Food Saf. 16:477-488, DOI:10.1111/1541-4337.12262

Rezaei M., A. Arzani, G. Saeidi and M. Karami (2017) Physiology of salinity tolerance in Bromus danthoniae genotypes originated from saline and non-saline areas of West Iran. Crop Pasture Sci. 68: 92–99.

Tohidi B., M. Rahimmalek and A. Arzani (2017) Essential oil composition, total phenolic, flavonoid contents, and antioxidant activity of Thymus species collected from different regions of Iran. Food Chem. 220: 153–161.

REVIEW PAPER: Arzani A. and M. Ashraf (2016) Smart engineering of genetic resources for enhanced salinity tolerance in crop plants. Crit. Rev. Plant Sci. 35: 146-189. Access to the full text (free HTML/pdf):

Saha P., N. Sade, A. Arzani, M.M.R. Wilhelmi, K.M. Coe, B. Li and E. Blumwald (2016) Effects of abiotic stress on physiological plasticity and water use of Setaria viridis (L.). Plant Sci. 251: 128-138.

Ebrahimi F., M. M. Majidi, A. Arzani and G. Mohammadi-Nejad (2016) Oil and seed yield stability in a worldwide collection of safflower under arid environments of Iran. Euphytica 212: 131–144. Access to the full text (pdf):

Ghorbani F. and A. Arzani (2016) Karyological studies in Triticum monococcum subsp. aegilopoides and  Aegilops cylindrica species  grown wild pairwise in west Iran.  Rostaniha (Iran J Bot) 16: 164-173.

Ghorbani F., A. Arzani and M.M. Poursiahbidi (2015) Meiotic instability in Aegilops cylindrica: a comparison with stable meiosis in Triticum monococcum subsp. aegilopoides. Caryologia 68: 101-108.

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Abbasi Z., MM Majidi, A Arzani, A Rajabi, P Mashayekhi, J. Bocianowski (2015) Association of SSR markers and morpho-physiological traits associated with salinity tolerance in sugar beet (Beta vulgaris L.). Euphytica 205: 785-797.Access to the full text (pdf):

Kiani R., A. Arzani and F. Habibi (2015) Physiology of salinity tolerance in Aegilops cylindrica. Acta Physiol. Plant. 37: 135. doi 10.1007/s11738-015-1881-0. Access to the full text (pdf):

Yeilaghi H., A. Arzani, M. Ghaderian (2015) Evaluating the contribution of ionic and agronomic components toward salinity tolerance in safflower. Agron. J. 107: 2205-2212.

Khaksar G., B. E. Sayed Tabatabae, A. Arzani, C. Ghobadi, E. Ebrahimie (2015) Functional analysis of a pomegranate (Punica granatum L.) MYB transcription factor involved in the regulation of anthocyanin biosynthesis. Iran J Biotech. 13: 17-25.

Bahrami, F., A. Arzani and V. Karimi (2014) Evaluation of yield-based drought tolerance indices for screening safflower (Carthamus tinctorius L.) genotypes. Agron. J. 106: 1219-1224.

Hodaei M., A. Arzani  and M. Rahimmalek (2014) Plasmon analysis in wheat alloplasmic lines using morphological and chloroplast microsatellite markers. Plant Syst. Evol. 300: 1137–1145. Access to the full text (pdf):


Journal Papers

Karimi S., G. Saeidi, A. Arzani, H. Amini and F. Bahrami (2014). Microsatellite markers variation and seed oil composition of some safflower genotypes differing in salt tolerance. J. Pure Appl. Microbiol. 9: 2077-2086.

Abbasi Z., A. Arzani, M. M. Majidi (2014) Evaluation of genetic diversity of sugar beet (Beta vulgaris L.) crossing parents using agro-morphological traits and molecular markers. J. Agric. Sci. Techn. 16: 1397-1411.
Salehi, M. and A. Arzani (2014) Evaluation of triticale genotypes for salt tolerance using physiological traits. Emir. J. Food Agric. 26: 277-283.

Amini, H., A. Arzani A. and M. Karimi (2014) Effect of water deficiency on seed quality and physiological traits of different safflower genotypes. Turk. J. Biol. 38: 271-282.

Arabbeigi M., A. Arzani, M.M. Majidi, R. Kiani, B.E. Seyed Tabatabaei, F. Habibi. (2014). Salinity tolerance of Aegilops cylindrica genotypes collected from hypersaline shore area of Uremia Salt Lake using physiological traits and SSR markers. Acta Physiol. Plant. 36: 2243-2251. Access to the full text (pdf):

Karimi S., A. Arzani and G. Saeidi (2014). Differential response of ion and osmolyte accumulation to salinity stress in salt-tolerant and salt-sensitive seedlings of safflower (Carthamus tinctorius L.). Crop Res. 15: 802-809.

Salehi, M. and A. Arzani (2013) Grain quality traits in triticale influenced by field salinity stress. Aust. J. Crop Sci. 7: 580-587.

Amini, H., A. Arzani A. and  F. Bahrami (2013) Seed yield and some physiological traits of safflower as affected by water deficit stress. Int J. Plant Prod. 7: 597-614.

Bahrami F., A. Arzani and H. Amini (2013) Leaf anatomical characteristics in safflower genotypes as affected by drought stress. Acta Biol. Szeged. 57: 39-42.

Abtahi, M. and A. Arzani (2013) Molecular and morphological assessment of genetic variability induced by gamma radiation in canola. J. Plant Mol. Breed. 1: 69-84.

Nazem, V. and A. Arzani (2013) Evaluation of morphological traits diversity in synthetic hexaploid wheat. J. Appl. Environ. Biol. Sci. 3: 20-28.

Golabadi, M., A. Arzani, S. A. M. Mirmohammadi Maibody, B. E. Sayed Tabatabaei and S. A. Mohammadi (2011). Identification of microsatellite markers linked with yield components under drought stress at terminal growth stages in durum wheat. Euphytica 177: 207-221. Access to the full text (pdf):

Genetic analysis of oil content and fatty acid composition in safflower (Carthamus tinctorius L.).

Golkar P., A. Arzani and A.M. Rezaei (2011). J. Amer. Oil Chem. Soc. 88: 975-982.

Genetic variation in safflower (Carthamus tinctorious L.) for seed quality-related traits and inter-simple sequence repeat (ISSR) markers.

Golkar P., A. Arzani and A.M. Rezaei (2011). Int. J. Mol. Sci. 12: 2664-2677

Evaluation of salinity tolerance in sorghum (Sorghum bicolor L.) using ion accumulation, proline and peroxidase criteria.

Bavei, V., B. Shiran and A. Arzani (2011) Plant Growth Regul. 64:275–285. Access to the full text (pdf):

Postulation of leaf rust resistance genes in Iranian wheat cultivars and breeding lines

Rafiei Boroujeni, F. A. Arzani, F. Afshari and M. Torabi (2011). Can. J. Plant Pathol. 33: 550-558.

Identification and inheritance of leaf rust resistance genes in the wheat cultivar Marvdasht

Rafiei, F., A. Arzani, F. Afshari and M. Torabi (2011)


Development and characterization of microsatellite markers for genomic analysis of yarrow (Achillea millefolium L.)

Rahimmalek M., B.E. Sayed Tabatabaei, A. Arzani and M. Khorrami (2011) Genes & Genomics 33: 475-482. Access to the full text (pdf):

Evaluation of triticale genotypes for terminal drought tolerance using physiological traits

Akbarian A., A. Arzani, M. Salehi and M. Salehi (2011) Indian J. Agic. Sci. 81: 1110–1115.

Morpho-physiological traits associated with terminal drought-stress tolerance in triticale and wheat

Lonbani, M. and A. Arzani (2011). Agron. Res. 9: 315-329.

Plant regeneration via somatic embryogenesis in three Iranian Cucumis melo L. genotypes

Moradmand, R., A. Arzani and G. Saeidi (2011) J. Crop Improv. 25: 183-190.

Inheritance of flower colour and spinelessness in safflower (Carthamus tinctorius L.).

Golkar, P., A. Arzani and A. M. Rezaei (2010) J. Genet. 89: 259-262.

Meiotic behavior of tetraploid wheats (Triticum turgidum L.) and their synthetic hexaploid wheat derivates influenced by meiotic restitution and heat stress

Rezaei, M., A. Arzani and B. E. Sayed-Tabatabaei (2010) J. Genet. 89: 401-407. Access to the full text (pdf):


Meiotic behavior of wild, synthetic and cultivated wheats

Arabbeigi, M., A. Arzani and G. Saeidi (2010) Cytologia 75: 169-175.

Genetic analysis of some agronomic traits in flax (Linum usitatissimum L.)

Mohammadi, A.A., G. Saeidi and A. Arzani (2010) Aust. J. Crop Sci. 4:343-352.


Essential oil variation in Thymus daenensis subsp. daenensis Cleak populations

Bahreininejad, B., M. Mirza and A. Arzani (2010) J. Essent. Oil Res. 22: 48-52.

Evaluation of salinity tolerance in rice genotypes

Mohammadi-Nejad, G., R.K. Singh, A. Arzani, A.M. Rezaie, H. Sabourid and G.B. Gregorio (2010) Int. J. Plant Prod. 4: 199-208

Assessment of genetic diversity among and within Achillea species using amplified fragment length polymorphism (AFLP).

Rahimmalek, M., B.E.Sayed Tabatabaei, A. Arzani and N. Etemadi, (2009). Biochem. Syst. Ecol. 37: 354–361.

Essential oil variation among and within six Achillea species transferred from different ecological regions in Iran to the field conditions.

Rahimmalek, M., B.E.Sayed Tabatabaei, N. Etemadi, S.A. H. Goli, A. Arzani and H. Zeinali (2009).Ind. Crops Prod. 29: 348-


Effects of irrigation water salinity and leaching fraction on yield and evapotranspiration in spring wheat

Heidarpour, M., B. Mostafazadeh-Fard, A. Arzani, A. Aghakhani and M. Feizi (2009) Commun. Soil Sci. Plant Anal. 40: 2521-2535. Access to the full text (pdf):


Study of genetic diversity in safflower genotypes using agro-morphological traits and RAPD markers.

Amini F., G. Saeidi and A. Arzani (2008).Access to the full text (pdf):


REVIEW PAPER: Improving salinity tolerance in crop plants: a biotechnological view

Arzani A. (2008) In Vitro Cell. Dev. Biol.-Plant (2008) 44:373–383


Assessment of rice genotypes for salt tolerance using microsatellite markers associated with the saltol/QTL

Mohammadi-Nejad G., A. Arzani, A. M. Rezai, R.K. Singh and G. B. Gregorio (2008) Afr. J. Biotech. 7: 730-736.


Characterization of leaf rust resistance genes in seedlings of wheat cultivars.

Rafiei, F., A. Arzani, F. Afshari and M. Torabi (2007)
Genet. Breed. 36: 19-27.

Iron and magnesium concentrations of mint accessions (Mentha spp.).

Arzani, A., H. Zeinali and K. Razmjo (2007)


Assessment of drought tolerance in segregating populations in durum wheat

Golabadi, M., A. Arzani and S.A.M. Mirmohammadi Maibody (2006) Afr. J. Agric. Res. 1: 162-171.

Evaluation of salt-tolerant genotypes of durum wheat derived from in vitro and field

Houshmand, S., A. Arzani and S.A.M. Maibody and M. Feizi (2005)
experiments. Field Crops Res. 91: 345-354.

Assessment of genetic diversity in bread wheat genotypes for tolerance to drought using AFLPs and agronomic traits.  Access to the full text (pdf):

Esmaeilzadeh Moghaddam, M., R.M. Trethowan, H.M. William, A. Rezai, A. Arzani and A.F. Mirlohi (2005)

Evaluation of diversity in wild relatives of wheat

Arzani, A., M.R. Khalighi, B. Shiran and N. Kharazian (2005) Czech J. Genet. Plant Breed. 41: 112-117

Evaluation of variation among durum wheat F3 families for grain yield and its componenets under normal and water-stress field conditions.

Golabadi, M., A. Arzani and S.M. Maibody (2005) Czech J. Genet. Plant Breed. 41: 263-267.

Essential oil composition of Iranian mints (Mentha spp.)

Zeinali, H., A. Arzani, K. Razmjo and M.B. Rezaee (2005). J. Essent. Oil Res. 17: 156-159.

DNA and morphological markers for a Russian wheat aphid resistance gene

Arzani, A., J. Peng and N.L.V. Lapitan (2004) Euphytica 139: 167-172.   Access to the full text (pdf):

Diversity among Iranian mints in relation to yield and mineral content

Zeinali, H., K. Razmjo and A. Arzani (2003) Commun. Soil Plant Anal. 34: 2217-2233.

Androgenetic response of heterozygous triticale populations using a greenhouse hydroponic system

Arzani, A. and N.L. Darvey (2002) Euphytica 127: 53-60.

Comparison of doubled haploid lines and their mid-generation progenitors in forage and dual-purpose triticales under greenhouse hydroponic conditions

Arzani, A. and N.L. Darvey (2002) Euphytica 126: 219-225.Access to the full text (pdf):

Grain yield performance of durum wheat germplasm under Iranian dryland and irrigated field conditions

Arzani, A. (2002) Sabrao J. Breed. Genet. 34: 9-18.

The effect of colchicine on triticale anther-derived plants: microspore pre-treatment and haploid-plant treatment using a hydroponic recovery system

Arzani, A. and N.L. Darvey (2001) Euphytica 122: 235-242.  Access to the full text (pdf):

Quantitative genetic analysis of forage and dual purpose characteristics of triticale using doubled haploid lines

Arzani, A. and N. L. Darvey (2001) Sabrao J. Breed. Genet. 33: 87-98.

Response of durum wheat cultivars to immature embryo culture, callus induction and in vitro salt stress

Arzani A. and S.S. Mirodjagh. (1999) Plant Cell Tiss. Org Cult. 58: 67-72.


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